Warning: SPOILERS for Star Trek: Lower Decks Season 5, Episode 9 - “Fission Quest"Star Trek: Lower Decksjust delivered the biggestStar Trekcrossover episode of all time, bringing back a jaw-dropping six legacy characters from variousStar Trekseries, along with the original actors voicing five of the characters.Star Trek: Lower Decks' final season on Paramount+ has centered on the USS Cerritos closing fissures in space that lead to alternateStar Trektimelines. However, another starship is assigned to repairing the multiverse, andit’s crewed by a host of alternate realityStar Treklegends.

Led by Captain William Boimler (Jack Quaid), the transporter clone of Lt. Bradward Boimler who faked his death and joined Section 31, the Defiant Class starship Anaximander has chased a vessel called the Beagle across countless quantum realities. Captain Boimler’s starship picks up several of thebest Star Trekcharactersfrom alternate realities to protect the multiverse. InStar Trek: Lower Decksseason 5, episode 9, “Fission Quest,” this includes an alternate reality version of Lt. Beckett Mariner (Tawny Newsome), who is a Starfleet engineer.

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Here are the iconicStar Trekcharacters who have returned inStar Trek: Lower Decks' penultimate episode.

T’Pol

Voiced By Jolene From Star Trek: Enterprise

T’Pol might be the biggest jaw-dropper among theStar Treklegends who have made a comeback inStar Trek: Lower Decksseason 5, episode 9, “Fissure Quest.” Jolene Blalock played Subcommander T’Pol for all four seasons ofStar Trek: Enterprise. In the interim years, however, Jolene got married, started a family, and left acting. Although she appeared in Star Trek Day 2021,it was widely believed Jolene leftStar Trekbehind. Incredibly,Star Trek: Lower Deckshas brought Jolene (credited with no surname) back to play T’Pol.

Star Trek: Enterprise Cast & Character Guide

Star Trek: Enterprise introduced new faces to the prequel series set a century before the events of Star Trek: The Original Series.

Star Trek: Lower Decks' alternate reality version of T’Pol has many similarities to the Prime Universe’s Vulcan fromStar Trek: Enterprise.The alt T’Pol was married to Trip Tucker (Connor Trinneer) for decadesbecause her Trip didn’t die as he did inStar Trek: Enterprise’s series finale. Being married to Trip gave T’Pol particular insight into humans, and Trills, as the Vulcan is friends with the surly Curzon Dax (Fred Tatasciore). T’Pol also recalls her version of Captain Jonathan Archer (Scott Bakula) was a"big Starfleet nerd,“which goes to show some things are consistent in any reality,

Bashir Lower Decks

Harry Kim

Voiced By Garrett Wang From Star Trek: Voyager

Garrett Wang returns asStar Trek: Voyager’s eternal ensign, Harry Kim, but with aStar Trek: Lower Deckstwist:Most of the Anaximander’s crew is comprised of alt universe Harry Kims picked up from multiple quantum realities. And, except for oneLieutenant Harry Kim, all of the other Harrys are still ensigns. Of course, Garrett Wang played Ensign Harry Kim for all seven seasons ofStar Trek: Voyager.

A plan for Garrett Wang to appear as Captain Harry Kim inStar Trek: Picardseason 3 didn’t happen due to budgetary issues.

Alexander Siddig as Julian Bashir in Our Man Bashir and Doctor Bashir, I Presume?

Star Trek: Lower Decksseason 5, episode 9 delivers another twist asLieutenant Harry Kim turns out to be the episode’s villain. Seething over how the gaggle of Ensign Kims are"disrespected,“Lieutenant Kim goes rogue and puts the entire multiverse in jeopardy.Star Trek: Lower Deckstells a great gag about how it seems to be Harry Kim’s cosmic destiny to remain thelowest-ranked Starfleet Officerin nearly every reality, and the one Harry Kim who rose in rank ends up being a threat to the multiverse.

Dr. Julian Bashir

Voiced By Alexander Siddig From Star Trek: Deep Space Nine

One of the most requestedStar Trek: Deep Space Ninecharacters to make a comeback, Alexander Siddig finally reprises Dr. Julian Bashir, but with a multiverse twist:Star Trek: Lower Decks' version of Bashir is an Emergency Medical Hologram.InStar Trek: Deep Space Nineseason 5, episode 16, “Doctor Bashir, I Presume?”, Julian was meant to be the basis for a new version of the EMH until it was discovered that the good doctor underwent illegal genetic modifications as a child.

10 Best Dr. Bashir Star Trek DS9 Episodes

Julian Bashir’s an augment, a doctor, and a secret agent, and his best Star Trek: Deep Space Nine episodes encapsulate these aspects of his character.

Star Trek: Lower Decks' holographic Dr. Bashir may or may not have been based on an augment in his universe, butJulian is in a relationship with Elim Garak (Andrew Robinson) from a different reality.Lower Decks’Bashir and Garak love match pays off an unfulfilled story that was only hinted at inStar Trek: Deep Space Nine.It’s also savvy thatStar Trek: Lower Decksplaces Dr. Bashir on a Section 31 starship whenDeep Space Nine’s Bashir was sought after and briefly became an unwilling operative of Section 31.

Lily Sloane Lower Decks

Elim Garak

Voiced By Andrew Robinson From Star Trek: Deep Space Nine

There’s no mistaking the smooth tone of Andrew Robinson as the voice of Elim Garak inStar Trek: Lower Decks.Garak is one ofStar Trek: Deep Space Nine’s most ingenious creations: a former Cardassian spy for the Obsidian Order who was exiled and set himself up as an innocent tailor on the DS9 space station. Eventually, even Captain Benjamin Sisko (Avery Brooks) came to rely on Garak’s skills and subterfuge. ButGarak’s most important Starfleet connection is his friendship with Dr. Julian Bashir.

OnStar Trek: Deep Space Nine, Tora Ziyal (Melanie Smith), the daughter of Gul Dukat (Marc Alaimo), was infatuated with Garak before her death.

Curzon Dax Lower Decks

Star Trek: Lower Decks season 5, episode 9 delivers the Bashir and Garak love story thatStar Trek: Deep Space Nineonly hinted at.Lower Decks' alt reality Garak and Bashir are in love, although they bicker over whose universe they will ultimately live in.Lower Decks’Garak is also a doctor, but he retains his lethal skillset as a Cardassian spy.But Garak’s greatest talent is infuriating Julian, which is his way of having a little fun while the multiverse is in jeopardy.

Lily Sloane

Voiced By Alfre Woodard From Star Trek: First Contact

Perhaps the most unexpectedStar Treklegacy actor to return inStar Trek: Lower Decksseason 5, episode 9 is Alfre Woodard, who reprisesLily Sloane fromStar Trek: First Contact.Lily worked alongside Dr. Zephram Cochrane (James Cromwell) to develop warp drive, which was installed on their starship dubbed the Phoenix. However, while Cochrane achieved Mankind’s first warp flight with crew members from the USS Enterprise-E, Lily was trapped on the starship fighting the Borg alongside Captain Jean-Luc Picard (Patrick Stewart).

Lily is an explorer who is fascinated with the human condition.

Jadzia Dax (Terry Farrell on DS9) and Boimler (Jack Quaid) in live-action on Strange New Worlds

Star Trek: Lower Decks' alt reality Lily Sloane has never heard of warp drive. This Lily Sloane is the Captain of the Beagle, a starship that, instead of warp drive,uses technology Lily and Cochrane created that can cross into different quantum realities.Lily is an explorer who is fascinated with the human condition. Sloane’s voyages throughout the multiverse aren’t about exploring new worlds but learning about human potential by studying Mankind in every reality.

Curzon Dax

Voiced By Fred Tatasciore From Star Trek: Lower Decks

Curzon Dax is the onlyStar Treklegacy character inLower Decks' penultimate episode who isn’t voiced by the original actor. InStar Trek: Deep Space Nine, Curzon was played by Frank Owen Smith, who had no dialogue, and by the late Rene Auberjonois. Fred Tatasciore, who voices Lt. Shax, among otherStar Trek: Lower Deckscharacters, steps in to playthe legendary Curzon Dax, who was the seventh Trill to host the Dax symbiont before Lt. Commander Jadzia Dax(Terry Farrell) and Lt. Ezri Dax (Nicole de Boer), Curzon was also Captain Sisko’s mentor.

After 26 Years, Star Trek Reveals A Dax DS9 Love Interest & It’s A Shocker

Star Trek: Lower Decks finally revealed what Gallamites look like 26 years after Lt. Commander Jadzia Dax dated one on Deep Space Nine.

Like the Prime Universe Curzon Dax,Star Trek: Lower Decks' alt reality Curzon must also have previously been Federation Ambassador to the Klingon Empire, judging from the bat’leth he hefts around.Curzon also has a healthy dislike for Vulcans in general,and T’Pol, in particular, and Dax is stunned that T’Pol has always considered him a friend.Star Trek: Lower Deckscleverly brings Curzon Dax back to life so that audiences can see him in action like never before.

Star Trek: Lower Decks Season 5 Official Poster